Right forelimb of specimen "Sue" The shoulder girdle was more time than your entire forelimb. The shoulder blade experienced a narrow shaft but was extremely expanded at its upper finish.
Paul turned down the objections lifted by critics, insisting that they're unwilling to contemplate that Tyrannosaurus could signify more than one species.[83] In a subsequent paper awaiting publication, Paul preserved the summary that Tyrannosaurus is made of three species. He identified the criticism on the study naming T. imperator and T. regina only focused on two of the characteristics made use of to differentiate The 2 new species (the number of little incisiform enamel and femur robustness), while the original review also compared the robustness of other bones also (the maxilla, dentary, humerus, ilium and metatarsals).
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A study by Grant R. Hurlburt, Ryan C. Ridgely and Lawrence Witmer acquired estimates for Encephalization Quotients (EQs), depending on reptiles and birds, together with estimates for the ratio of cerebrum to brain mass. The review concluded that Tyrannosaurus experienced the reasonably largest brain of all adult non-avian dinosaurs except sure little maniraptoriforms (Bambiraptor, Troodon and Ornithomimus). The study identified that Tyrannosaurus's relative brain measurement was continue to within the variety of recent reptiles, being at most 2 regular deviations previously mentioned the signify of non-avian reptile EQs.
A discussion exists, KS Quik even so, about irrespective of whether Tyrannosaurus was primarily a predator or possibly a pure scavenger. The controversy originated in the 1917 examine by Lambe which argued that giant theropods were being pure scavengers for the reason that Gorgosaurus enamel confirmed barely any use.
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